From the Illustrator help pages

The target icon indicates whether an item in the layer hierarchy has any appearance attributes and whether it is targeted

The filled in icon means the layer has appearance attributes beyond a single fill and a single stroke.

You can read more here: helpx.adobe.com/illustrator/using/appearance-attributes.html

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: Can somone explain the layer target icon? In the Illustrator layer pallete, somtimes the layer target icon is filled in, and somtimes not. Can somone explain to me what the difference is?

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